Animalia > Chordata > Aves > Charadriiformes > Alcidae > Cepphus > Cepphus grylle

Cepphus grylle (Black Guillemot)

Language: French

Wikipedia Abstract

The black guillemot or tystie (Cepphus grylle) is a medium-sized alcid. The genus name Cepphus is from Ancient Greek kepphos, a pale waterbird mentioned by Greek authors including Aristotle. The species name grylle was the local dialect name for this bird in Gotland at the time of Linnaeus's visit there in 1741. The English word "guillemot" is from French guillemot probably derived from Guillaume, "William". There are five listed subspecies of the black guillemot: These birds often overwinter in their breeding areas, moving to open waters if necessary, but usually not migrating very far south.
View Wikipedia Record: Cepphus grylle


EDGE Analysis

Uniqueness Scale: Similiar (0) 
 Unique (100)
Uniqueness & Vulnerability Scale: Similiar & Secure (0) 
 Unique & Vulnerable (100)
ED Score: 8.62204
EDGE Score: 2.26406


Adult Weight [2]  428 grams
Birth Weight [2]  33 grams
Female Maturity [2]  3 years
Male Maturity [2]  3 years
Breeding Habitat [3]  Coastal cliffs and islands, Coastal marine
Wintering Geography [3]  Atlantic Coast
Wintering Habitat [3]  Coastal marine
Diet [4]  Carnivore (Invertebrates), Piscivore
Diet - Fish [4]  60 %
Diet - Invertibrates [4]  40 %
Forages - Underwater [4]  100 %
Clutch Size [2]  2
Clutches / Year [2]  1
Global Population (2017 est.) [3]  750,000
Incubation [2]  25 days
Maximum Longevity [2]  26 years
Migration [1]  Intracontinental
Water Biome [1]  Coastal
Wing Span [5]  22 inches (.55 m)


Name Countries Ecozone Biome Species Report Climate Land
Arctic desert Norway, Russia Palearctic Tundra    
Kamchatka-Kurile meadows and sparse forests Russia Palearctic Boreal Forests/Taiga
Northwest Russian-Novaya Zemlya tundra Russia Palearctic Tundra
Novosibirsk Islands arctic desert Russia Palearctic Tundra
Scandinavian and Russian taiga Norway, Sweden, Finland, Russia Palearctic Boreal Forests/Taiga
South Sakhalin-Kurile mixed forests Russia Palearctic Temperate Broadleaf and Mixed Forests    
Taimyr-Central Siberian tundra Russia Palearctic Tundra
Wrangel Island arctic desert Russia Palearctic Tundra    

Protected Areas

Name IUCN Category Area acres Location Species Website Climate Land Use
Acadia National Park II 35996 Maine, United States
Adlergrund 57815 Germany    
Alaska Maritime National Wildlife Refuge Ib 2347132 Alaska, United States
Arctic National Wildlife Refuge Ib 12402936 Alaska, United States
Ascrib, Isay and Dunvegan 6388 Scotland, United Kingdom
Beara Peninsula SPA 6493 Ireland  
Björkör 13062 Finland    
Björns skärgård 2769 Sweden    
Bonden 979 Sweden    
Cape Breton Highlands National Park II 234333 Nova Scotia, Canada  
Cape Cod National Seashore II 21724 Massachusetts, United States
Cape Wrath 2509 Scotland, United Kingdom  
Clare Island SPA 2486 Ireland  
Cliffs of Moher SPA 2160 Ireland  
Coll Machair 2104 Scotland, United Kingdom  
Dee Estuary/ Aber Dyfrdwy 39057 England/Wales, United Kingdom
Deenish Island and Scariff Island SPA 2090 Ireland  
Durness 2997 Scotland, United Kingdom  
East Caithness Cliffs 1094 Scotland, United Kingdom
Eileanan agus Sgeiran Lios mór 2816 Scotland, United Kingdom  
Fair Isle 1387 Scotland, United Kingdom  
Faray and Holm of Faray 1941 Scotland, United Kingdom  
Firth of Lorn 51830 Scotland, United Kingdom
Forillon National Park II 61010 Quebec, Canada  
Fundy National Park II 52716 New Brunswick, Canada
Gros Morne National Park II 476632 Newfoundland, Canada
Helvick Head to Ballyquin SPA 1939 Ireland  
Hoburgs bank 303019 Sweden  
Horn Head to Fanad Head SPA 6006 Ireland    
Hoy 23474 Scotland, United Kingdom
Itäisen Suomenlahden saaristo ja vedet 236302 Finland    
Iveragh Peninsula SPA 8620 Ireland  
Ivvavik National Park II 2382752 Yukon, Canada
Kejimkujik Seaside Adjunct National Park II 4970 Nova Scotia, Canada
Kirkkonummen saaristo (SPA) 35173 Finland  
Kokkolan saaristo (SPA) 36322 Finland  
Koster 2884 Sweden  
Kristiinankaupungin saaristo 19914 Finland  
Kura kurgu 476839 Estonia  
Lambay Island SPA 1482 Ireland  
Laplandskiy Biosphere Reserve 757114 Russia  
Lawica Slupska 197809 Poland    
Loch Maree Complex 38882 Scotland, United Kingdom
Loch nam Madadh 5734 Scotland, United Kingdom
Lochs Duich, Long and Alsh Reefs 5883 Scotland, United Kingdom  
Luce Bay and Sands 120487 Scotland, United Kingdom
Luodon saaristo 35731 Finland    
Luvian saaristo 18785 Finland  
Malmöfjord 1728 Sweden  
Merenkurkun saaristo 316695 Finland    
Mid-Clare Coast SPA 11467 Ireland  
Mingan Archipelago National Park Reserve II 20461 Quebec, Canada
Missjö 4529 Sweden  
Monach Islands 9011 Scotland, United Kingdom    
Moray Firth 373987 Scotland, United Kingdom  
Mousa 1311 Scotland, United Kingdom
Norra Midsjöbanken 243037 Sweden      
North Rona 1553 Scotland, United Kingdom    
North-east Greenland National Park II 237590314 Greenland
Oldshoremore and Sandwood 1096 Scotland, United Kingdom    
Pakri 50588 Estonia  
Papa Stour 5132 Scotland, United Kingdom  
Pernajanlahtien ja Pernajan saariston merensuojelualue 162534 Finland  
Perämeren saaret 17633 Finland  
Polar Bear Provincial Park 5502026 Ontario, Canada
Przybrzezne wody Baltyku 480933 Poland    
Queen Maud Gulf Bird Sanctuary Migratory Bird Sanctuary IV 15335601 Nunavut, Canada
Raahen saaristo 5535 Finland    
Rahjan saaristo 20710 Finland    
Ronas Hill – North Roe 12110 Scotland, United Kingdom  
Rum 26775 Scotland, United Kingdom
Rödkallen-SörÄspen 17514 Sweden    
SPA Pommersche Bucht 495241 Germany    
Saaristomeri 147873 Finland    
Saguenay - St. Lawrence Marine Park National Marine Conservation Area II 310822 Canada
Sanday 27112 Scotland, United Kingdom    
Seksmiilarin saaristo 42581 Finland  
Sheep's Head to Toe Head SPA 6181 Ireland    
Sirmilik National Park II 5475284 Canada
South-East Islay Skerries 3702 Scotland, United Kingdom
Stora Nassa 7286 Sweden  
Sullom Voe 6668 Scotland, United Kingdom
Sälöfjorden 7089 Sweden  
Söderskärin ja Långörenin saaristo 45020 Finland  
Tagamõisa 33316 Estonia    
Tanumskusten 20446 Sweden  
Terra Nova National Park IV 125894 Newfoundland, Canada
Tingon 1407 Scotland, United Kingdom    
Tory Island SPA 1501 Ireland    
Treshnish Isles 4850 Scotland, United Kingdom
Uudenkaarlepyyn saaristo 7932 Finland    
Uudenkaupungin saaristo 140830 Finland  
Virginia Coast Biosphere Reserve 33386 Virginia, United States  
Vrångöskärgården 17306 Sweden  
Västerbådan, Lågagrundet 3951 Sweden    
Wapusk National Park II 2614324 Manitoba, Canada
West Donegal Coast SPA 8347 Ireland    
West Estonian Archipelago Biosphere Reserve 3854844 Estonia  
Yell Sound Coast 3807 Scotland, United Kingdom
Zatoka Pomorska 770666 Poland    

Important Bird Areas

BirdLife Species Factsheet: View Factsheet
Name Location  IBA Criteria   Website   Climate   Land Use 
Île aux Basques et Les Razades Canada A4i
Ædey Iceland A4i, A4iii, B1i, B1ii, B2  
Bezymyannaya and Gribovaya Bays and adjoining waters Russia (European) A1, A4i, A4ii, A4iii, B1i, B1ii, B3
Björn archipelago Sweden A4i, B1i, B1ii, B2, B3, C2, C3, C6  
Bjørnøya (Bear Island) Svalbard and Jan Mayen Islands (to Norway) A3, A4i, A4ii, A4iii, B1i, B1ii, B2  
Blasket Islands Ireland A4ii, A4iii, B1ii, B2, B3, C2, C3, C4, C6    
Bonden island Sweden B1ii, B2, B3, C3  
Bordoy Faroe Islands (to Denmark) A4ii, B1ii, B2  
Bråviken: Hävringe Sweden A4i, A4iii, B1i, B1ii, B2, B3, C2, C3, C4, C6
Breidafjördur Iceland A4i, A4ii, A4iii, B1i, B1ii, B2, B3
Caithness Cliffs United Kingdom A4i, A4ii, A4iii, B1i, B1ii, B2, B3, C2, C3, C4
Central Polish coastal waters Poland C3, C4  
Clare Island cliffs Ireland B2
Digges Sound Canada A4ii, A4iii
Duvillaun Islands Ireland B1i, B2, B3, C2  
Eckerö and Hammarland archipelago Finland A4i, B1i, B1ii, B2, B3, C2, C3, C6  
Eysturoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2  
Faray and Holm of Faray United Kingdom B2, B3  
Forlandet National Park Svalbard and Jan Mayen Islands (to Norway) A4iii, B1i, B2, B3
Froan Norway A1, A4i, B1i, B1ii, B2, B3  
Fugloy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3  
Gigha Island and Islets United Kingdom B1ii, B2
Hermaness, Saxa Vord and Valla Field United Kingdom A4ii, A4iii, B1ii, B2, B3, C3, C4  
Hestur Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2
Hirsholmene Denmark A4i, B1i, B1ii, B2, B3, C6  
Hoburgs Bank Sweden A4i, B1i, B1ii, C3
Holmöarna archipelago Sweden A4i, A4ii, B1i, B1ii, B2, C2, C3, C6
Hopen island Svalbard and Jan Mayen Islands (to Norway) A4i, A4ii, A4iii, B1i, B1ii, B2  
Howth Head Ireland B2, C7
Hyllekrog-Rødsand and Fehmarn Belt Denmark A4i, A4iii, B1i, B1ii, C2, C3, C4
Inishtrahull Ireland B2, B3, C7  
Irbe strait Latvia A4i, A4ii, A4iii, B1i, B1ii
Irbe strait Estonia A1, A4i, A4iii, B1i, B1ii, C1, C2, C3, C4, C6
Islands and waters south and west of Upernavik town Greenland (to Denmark) A4iii, B1ii, B2
Itäinen Suomenlahti National Park Finland B1i, B1ii, B2, B3, C2, C3, C6
Jan Mayen island Svalbard and Jan Mayen Islands (to Norway) A4ii, A4iii, B1ii, B2  
Kalsoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2  
Kandalaksha Bay Russia (European) A1, A4i, A4iii, B1i, B1ii, B2
Koltur Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2
Korppoo and Nauvo southern archipelago Finland A4i, A4iii, B1i, B1ii, B2, B3, C3, C4  
Kunoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2  
Kuril islands (between Urup and Paramushir) Russia (Asian) A1, A4i, A4ii, A4iii  
Loppa Norway A4iii, B1ii, B2, B3  
Lovunden Norway A1, A4iii, B1ii, B2  
Merenkurkku archipelago Finland A4i, A4iii, B1i, B1ii, B1iv, B2, B3, C2, C3, C4, C6  
Monach Islands United Kingdom B1i, B1ii, B2, C2    
Mulklobb Finland B1ii, B2, C3  
Mykines and Mykineshólmur Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3    
Nólsoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2
North Ronaldsay Coast United Kingdom C7    
North Westray Coast United Kingdom B1ii, B2, C2, C3, C6  
North-west Spitsbergen National Park Svalbard and Jan Mayen Islands (to Norway) A4i, A4iii, B1i, B2, B3  
Northeast James Bay Coast Canada A4i, A4ii, A4iii
Northern Midsjö Bank Sweden A4i, B1i, B1ii, C3    
Onega Bay of the White Sea Russia (European) A4i, B1i, B1ii, B2, B3  
Outer Stockholm archipelago Sweden A4i, A4ii, A4iii, B1i, B1ii, B2, B3, C2, C3, C4, C6
Papa Westray (North Hill and Holm) United Kingdom B1ii, B2, C2, C3, C6  
Péninsule de Forillon Canada A4iii  
Pernaja outer archipelago Finland B1i, B1ii, B2, B3, C2, C3, C6
Pomeranian bay Poland C2, C3, C4  
Porvoo outer archipelago Finland A4i, B1i, B1ii, B2, C2, C6
Prince Leopold Island Canada A4ii, A4iii  
Rahja archipelago Finland B1i, B2, B3, C3  
Rödkallen-SörÄspen Sweden B1i, B1ii, B2, C2, C6  
Røst Norway A4i, A4ii, A4iii, B1i, B1ii, B2, B3    
Rockabill Ireland A4i, B1i, B2, C2, C6
Ronne Banke Denmark A4i, A4ii, A4iii, B1i, B1ii, C3, C4  
Sandoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3
Sør-Fugløy Norway A1, A4ii, A4iii, B1ii, B2, B3    
Scapa Flow United Kingdom A4i, B1i, B1ii, B2, C2, C3, C6
Skúvoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3
Sklinna Norway A4i, B1i, B1ii, B2, B3    
Slupsk bank Poland C3, C4  
Smøla archipelago Norway A1, A4i, B1i, B1ii, B2  
South-eastern Stronsay United Kingdom B2  
Southern Midsjö Bank Sweden B1ii    
Stóra Dímun Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2
Streymoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3  
Suduroy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3
Svínoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2  
Tammisaari and Inkoo western archipelago Finland A1, A4i, B1i, B2, B3, C1, C2, C6
The Archipelago of Nothern Hälsingland Sweden B1i, B1ii, B2, B3, C3, C6  
Vágar Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2, B3  
Väderöarna Sweden A4i, B1i, B2, B3, C3
Vidoy Faroe Islands (to Denmark) A4ii, A4iii, B1ii, B2  

Prey / Diet

Ammodytes americanus (inshore sand lance)[6]
Ammodytes hexapterus (Stout sand lance)[6]
Ammodytes marinus (sand eel)[6]
Boreogadus saida (Polar cod)[7]
Eleginus gracilis (Wachna cod)[6]
Enchelyopus cimbrius (Rockling)[6]
Eumesogrammus praecisus (Fourline snakeblenny)[8]
Mallotus villosus (Capelin)[9]
Microgadus tomcod (Frostfish)[6]
Myoxocephalus scorpius (Short-spined sea scorpion)[7]
Pholis gunnellus (Tissy)[6]
Stichaeus punctatus (Arctic shanny)[8]

Prey / Diet Overlap

Competing SpeciesCommon Prey Count
Alca torda (Razorbill)1
Amblyraja hyperborea (Arctic skate)1
Amblyraja radiata (Starry ray)1
Artediellus atlanticus (Atlantic hookear)1
Balaenoptera acutorostrata (Minke Whale)1
Balaenoptera borealis (Sei Whale)1
Balaenoptera physalus (Fin Whale)1
Boreogadus saida (Polar cod)1
Brachyramphus brevirostris (Kittlitz's Murrelet)1
Brachyramphus marmoratus (Marbled Murrelet)1
Careproctus reinhardti (Longfin seasnail)1
Cystophora cristata (Hooded Seal)1
Delphinapterus leucas (Beluga)2
Erignathus barbatus (Bearded Seal)2
Fratercula arctica (Atlantic Puffin)1
Fratercula cirrhata (Tufted Puffin)1
Fratercula corniculata (Horned Puffin)1
Fulmarus glacialis (Northern Fulmar)1
Gadus morhua (rock cod)1
Halichoerus grypus (Gray Seal)1
Hexagrammos stelleri (Greenling)1
Hippoglossoides platessoides (American dab)1
Histriophoca fasciata (Ribbon Seal)1
Lagenorhynchus albirostris (White-beaked Dolphin)1
Larus argentatus (Herring Gull)1
Larus canus (Mew Gull)1
Larus glaucescens (Glaucous-winged Gull)1
Larus marinus (Great Black-backed Gull)1
Liparis gibbus (Dusky snailfish)1
Lycodes reticulatus (Arctic eelpout)1
Malacoraja senta (Smooth skate)1
Megaptera novaeangliae (Humpback Whale)1
Melanogrammus aeglefinus (Smokie)1
Monodon monoceros (Narwhal)1
Morus bassanus (Northern Gannet)1
Orcinus orca (Killer Whale)1
Pagophila eburnea (Ivory Gull)1
Pagophilus groenlandicus (Harp Seal)2
Phalacrocorax aristotelis (Shag)1
Phalacrocorax auritus (Double-crested Cormorant)1
Phalacrocorax carbo (Great Cormorant)2
Phoca largha (Spotted Seal)1
Phocoena phocoena (Harbor Porpoise)1
Pollachius virens (Sillock)1
Pusa hispida (Ringed Seal)2
Reinhardtius hippoglossoides (Turbot)1
Rissa tridactyla (Black-legged Kittiwake)2
Sebastes mentella (Redfish)1
Stercorarius parasiticus (Parasitic Jaeger)1
Triglops murrayi (Moustache sculpin)1
Uria aalge (Common Murre)1
Uria lomvia (Thick-billed Murre)2


Haliaeetus leucocephalus (Bald Eagle)[6]


Parasitized by 
Alcataenia campylacantha[10]
Ceratophyllus garei[11]
Eucoleus contortus <Unverified Name>[10]

Institutions (Zoos, etc.)

Institution Infraspecies / Breed 
Biodome de Montreal
Jurong Birdpark, Singapore
National Aquarium in Baltimore Inc

Range Map

North America;



Attributes / relations provided by
1Myers, P., R. Espinosa, C. S. Parr, T. Jones, G. S. Hammond, and T. A. Dewey. 2006. The Animal Diversity Web (online). Accessed February 01, 2010 at
2de Magalhaes, J. P., and Costa, J. (2009) A database of vertebrate longevity records and their relation to other life-history traits. Journal of Evolutionary Biology 22(8):1770-1774
3Partners in Flight Avian Conservation Assessment Database, version 2017. Accessed on January 2018.
4Hamish Wilman, Jonathan Belmaker, Jennifer Simpson, Carolina de la Rosa, Marcelo M. Rivadeneira, and Walter Jetz. 2014. EltonTraits 1.0: Species-level foraging attributes of the world's birds and mammals. Ecology 95:2027
5British Trust for Ornithology
6Jorrit H. Poelen, James D. Simons and Chris J. Mungall. (2014). Global Biotic Interactions: An open infrastructure to share and analyze species-interaction datasets. Ecological Informatics.
7Notes on fishes in Hornsund fjord area (Spitsbergen), Jan Marcin WĘSLAWSKI and Wojciech KULIŃSKI, POLISH POLAR RESEARCH (POL. POLAR RES.) Vol. 10 No. 2 p. 241-250 (1989)
8"Diet and foraging ecology of Black Guillemots in northeastern Hudson Bay", D. K. Cairns, Canadian Journal of Zoology, 1987, 65(5): 1257-1263
9The role of capelin (Mallotus villosus) in the foodweb of the Barents Sea, A. V. Dolgov, ICES Journal of Marine Science, 59: 1034–1045. 2002
10Gibson, D. I., Bray, R. A., & Harris, E. A. (Compilers) (2005). Host-Parasite Database of the Natural History Museum, London
11International Flea Database
Ecoregions provided by World Wide Fund For Nature (WWF). WildFinder: Online database of species distributions, ver. 01.06 WWF WildFINDER
Protected Areas provided by Biological Inventories of the World's Protected Areas in cooperation between the Information Center for the Environment at the University of California, Davis and numerous collaborators.
Natura 2000, UK data: © Crown copyright and database right [2010] All rights reserved. Ordnance Survey Licence number 100017955
GBIF Global Biodiversity Information Facility
Images provided by Google Image Search
Abstract provided by DBpedia licensed under a Creative Commons License
Weather provided by NOAA METAR Data Access